Dr. Cheng is Associate Professor of Pathology, of Biochemistry & Molecular Biology, and of Pharmacology in the Jake Gittlen Cancer Research Foundation at the Penn State College of Medicine. He received a Bachelor's degree in Biochemical Sciences from Harvard University and an M.D. from New York University School of Medicine. He trained in Anatomic Pathology at Brigham & Women's Hospital in Boston and University of Washington, and while engaged in the latter, earned a Ph.D. in phage genetics at the Fred Hutchinson Cancer Research Institute and the University of Washington in Seattle. During his postdoctoral training at the University of Washington, Dr. Cheng studied mechanisms of mutation.