The scientific studies of sensory systems (including olfactory organs) are carried out during a century. Usually a preliminary theory of object is suggested and detailed in accordance with the new data. However only the strong criticized theories of smell reception more than thirty) are now in the theoretical part of the olfactory organ physiology. The theory is necessary for planning the further investigations of olfactory organ; revealing the mystery of the biological sensors to improve the artificial ones; the improvement of the methods to obtain new biological active substances (perfume, drugs, insecticides). These circumstances justify the attempt to suggest the theory of the olfactory receptor cell functioning.